誰 ___ このケーキを食べましたか。
The particle "が" (ga) marks the subject of a sentence. It's commonly used with interrogative words like "誰" (who), "何" (what), or "どれ" (which one) when the question is about identifying the subject performing the action.
助詞(じょし)「が」は文(ぶん)の主語(しゅご)を示(しめ)します。「誰(だれ)」「何(なに)」などの疑問詞(ぎもんし)と使(つか)い、動作(どうさ)の主(ぬし)を尋(たず)ねる際(さい)に使(つか)われます。例(れい)：「何(なに)が美味(おい)しいですか。」
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